169 lines
4.7 KiB
Python
169 lines
4.7 KiB
Python
|
"""Trophic levels"""
|
||
|
import networkx as nx
|
||
|
|
||
|
from networkx.utils import not_implemented_for
|
||
|
|
||
|
__all__ = ["trophic_levels", "trophic_differences", "trophic_incoherence_parameter"]
|
||
|
|
||
|
|
||
|
@not_implemented_for("undirected")
|
||
|
def trophic_levels(G, weight="weight"):
|
||
|
r"""Compute the trophic levels of nodes.
|
||
|
|
||
|
The trophic level of a node $i$ is
|
||
|
|
||
|
.. math::
|
||
|
|
||
|
s_i = 1 + \frac{1}{k^{in}_i} \sum_{j} a_{ij} s_j
|
||
|
|
||
|
where $k^{in}_i$ is the in-degree of i
|
||
|
|
||
|
.. math::
|
||
|
|
||
|
k^{in}_i = \sum_{j} a_{ij}
|
||
|
|
||
|
and nodes with $k^{in}_i = 0$ have $s_i = 1$ by convention.
|
||
|
|
||
|
These are calculated using the method outlined in Levine [1]_.
|
||
|
|
||
|
Parameters
|
||
|
----------
|
||
|
G : DiGraph
|
||
|
A directed networkx graph
|
||
|
|
||
|
Returns
|
||
|
-------
|
||
|
nodes : dict
|
||
|
Dictionary of nodes with trophic level as the vale.
|
||
|
|
||
|
References
|
||
|
----------
|
||
|
.. [1] Stephen Levine (1980) J. theor. Biol. 83, 195-207
|
||
|
"""
|
||
|
try:
|
||
|
import numpy as np
|
||
|
except ImportError as e:
|
||
|
raise ImportError("trophic_levels() requires NumPy: http://numpy.org/") from e
|
||
|
|
||
|
# find adjacency matrix
|
||
|
a = nx.adjacency_matrix(G, weight=weight).T.toarray()
|
||
|
|
||
|
# drop rows/columns where in-degree is zero
|
||
|
rowsum = np.sum(a, axis=1)
|
||
|
p = a[rowsum != 0][:, rowsum != 0]
|
||
|
# normalise so sum of in-degree weights is 1 along each row
|
||
|
p = p / rowsum[rowsum != 0][:, np.newaxis]
|
||
|
|
||
|
# calculate trophic levels
|
||
|
nn = p.shape[0]
|
||
|
i = np.eye(nn)
|
||
|
try:
|
||
|
n = np.linalg.inv(i - p)
|
||
|
except np.linalg.LinAlgError as err:
|
||
|
# LinAlgError is raised when there is a non-basal node
|
||
|
msg = (
|
||
|
"Trophic levels are only defined for graphs where every "
|
||
|
+ "node has a path from a basal node (basal nodes are nodes "
|
||
|
+ "with no incoming edges)."
|
||
|
)
|
||
|
raise nx.NetworkXError(msg) from err
|
||
|
y = n.sum(axis=1) + 1
|
||
|
|
||
|
levels = {}
|
||
|
|
||
|
# all nodes with in-degree zero have trophic level == 1
|
||
|
zero_node_ids = (node_id for node_id, degree in G.in_degree if degree == 0)
|
||
|
for node_id in zero_node_ids:
|
||
|
levels[node_id] = 1
|
||
|
|
||
|
# all other nodes have levels as calculated
|
||
|
nonzero_node_ids = (node_id for node_id, degree in G.in_degree if degree != 0)
|
||
|
for i, node_id in enumerate(nonzero_node_ids):
|
||
|
levels[node_id] = y[i]
|
||
|
|
||
|
return levels
|
||
|
|
||
|
|
||
|
@not_implemented_for("undirected")
|
||
|
def trophic_differences(G, weight="weight"):
|
||
|
r"""Compute the trophic differences of the edges of a directed graph.
|
||
|
|
||
|
The trophic difference $x_ij$ for each edge is defined in Johnson et al.
|
||
|
[1]_ as:
|
||
|
|
||
|
.. math::
|
||
|
x_ij = s_j - s_i
|
||
|
|
||
|
Where $s_i$ is the trophic level of node $i$.
|
||
|
|
||
|
Parameters
|
||
|
----------
|
||
|
G : DiGraph
|
||
|
A directed networkx graph
|
||
|
|
||
|
Returns
|
||
|
-------
|
||
|
diffs : dict
|
||
|
Dictionary of edges with trophic differences as the value.
|
||
|
|
||
|
References
|
||
|
----------
|
||
|
.. [1] Samuel Johnson, Virginia Dominguez-Garcia, Luca Donetti, Miguel A.
|
||
|
Munoz (2014) PNAS "Trophic coherence determines food-web stability"
|
||
|
"""
|
||
|
levels = trophic_levels(G, weight=weight)
|
||
|
diffs = {}
|
||
|
for u, v in G.edges:
|
||
|
diffs[(u, v)] = levels[v] - levels[u]
|
||
|
return diffs
|
||
|
|
||
|
|
||
|
@not_implemented_for("undirected")
|
||
|
def trophic_incoherence_parameter(G, weight="weight", cannibalism=False):
|
||
|
r"""Compute the trophic incoherence parameter of a graph.
|
||
|
|
||
|
Trophic coherence is defined as the homogeneity of the distribution of
|
||
|
trophic distances: the more similar, the more coherent. This is measured by
|
||
|
the standard deviation of the trophic differences and referred to as the
|
||
|
trophic incoherence parameter $q$ by [1].
|
||
|
|
||
|
Parameters
|
||
|
----------
|
||
|
G : DiGraph
|
||
|
A directed networkx graph
|
||
|
|
||
|
cannibalism: Boolean
|
||
|
If set to False, self edges are not considered in the calculation
|
||
|
|
||
|
Returns
|
||
|
-------
|
||
|
trophic_incoherence_parameter : float
|
||
|
The trophic coherence of a graph
|
||
|
|
||
|
References
|
||
|
----------
|
||
|
.. [1] Samuel Johnson, Virginia Dominguez-Garcia, Luca Donetti, Miguel A.
|
||
|
Munoz (2014) PNAS "Trophic coherence determines food-web stability"
|
||
|
"""
|
||
|
try:
|
||
|
import numpy as np
|
||
|
except ImportError as e:
|
||
|
raise ImportError(
|
||
|
"trophic_incoherence_parameter() requires NumPy: " "http://scipy.org/"
|
||
|
) from e
|
||
|
|
||
|
if cannibalism:
|
||
|
diffs = trophic_differences(G, weight=weight)
|
||
|
else:
|
||
|
# If no cannibalism, remove self-edges
|
||
|
self_loops = list(nx.selfloop_edges(G))
|
||
|
if self_loops:
|
||
|
# Make a copy so we do not change G's edges in memory
|
||
|
G_2 = G.copy()
|
||
|
G_2.remove_edges_from(self_loops)
|
||
|
else:
|
||
|
# Avoid copy otherwise
|
||
|
G_2 = G
|
||
|
diffs = trophic_differences(G_2, weight=weight)
|
||
|
return np.std(list(diffs.values()))
|